More about Elder Princess Shroob...
Games: Mario and Luigi: Partners in Time
 
Elder Princess Shroob is the leader of the Shroob invasion force that attacked the Mushroom Kingdom in the past, but she was not revealed until late in the adventure since she was trapped by Peach within the Cobalt Star when the princess first went back in time.  At that point her younger sister took command and the Marios never knew there was another Shroob leader while they unwittingly set about covering all the pieces of the Cobalt Star, not knowing that putting them altogether would release such a powerful foe.  EPS seems to have retained some consciousness while within the Cobalt Star.  Not only was she able to communicate with the Marios at one point, taking on the guise of a so-called Star Sprite, but she evidently learned English while imprisoned, as she communicated in that language although no other Shroob was shown to be able to do so.

EPS comes across as the power arm of her government, while her younger sister may have been more suited for leadership anyhow.  In battle she really throws her weight around and enjoys using tentacle attacks in addition to calling on her army's flying saucers to do all sorts of things.  In a straight-up fight she's quite difficult, but if you have any kind of inventory I think she can be managed, even though she is difficult to damage.  First her legs must be defeated to drop her crown into the field of play, then the crown needs to be taken out, and only then can her head, the true target, be damaged for a few turns until the other targets are revived.  With the defeat of EPS, the Shroob invasion seemed to be over for good, but EPS had one trick left.  After the battle her body morphed into a Shroob Mushroom that the Marios took back with them to their present time.  Bowser ate that Mushroom and EPS took control from the inside, making him incredibly powerful and able to attack rapidly.  Somehow the Marios defeated EPS in a way that doesn't quite make sense, Bowser was knocked out, and EPS presumably was destroyed for good.  Well, I doubt she was, but it's fun to pretend.

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